4.0 故事以高仓家三兄妹为中心而展开,高仓冠叶(木村昴 配音)、高仓晶马(木村良平 配音)兄弟俩和患有绝症的妹妹高仓阳毬( 荒川美穂 配音)。 有妹控情结的高仓兄弟俩都非常珍视阳毬,然而医生却让他们做好随时失去妹妹的准备。这天兄妹三人一同去水族馆游玩,可是一转眼兄弟二人发现妹妹不见了,随后他们在围观人群中央发现倒地不醒的阳毬。抢救无效被宣布死亡的阳毬竟然在停尸房里醒过来了!惊讶无比的兄弟俩把妹妹接回家,不久后他们收到了一份奇怪的包裹,打开后竟然发现里面有三只谜样企鹅。这三只企鹅憨态可掬,除了他们,外人都看不见这些企鹅。这一切究竟是怎么回事,命运的齿轮又会如何转动?
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